Property links: one layer follows another

A link makes one layer's number follow another layer's, so it moves without keyframes of its own.

Try this

  1. Press play. The blue shapes have no keyframes; each one follows another layer.
  2. Right-click a follower's linked Position field in Transform, choose Edit link…, and change Layer.
  3. To make one, right-click a Position field, choose Pick-whip to a layer, and click the leader.